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Rickettsiales

Rickettsias

Rickettsia rickettsii

Scientific classification

Kingdom:
Bacteria

Phylum:
Proteobacteria

Class:
Alphaproteobacteria

Subclass:
Rickettsidae

Order:
Rickettsiales
Gieszczkiewicz, 1939

Families

Rickettsiaceae
Midichloriaceae
Anaplasmataceae [1]

The Rickettsiales, also called rickettsias, are an order of small proteobacteria. Most of those described survive only as endosymbionts of other cells. Some are notable pathogens, including Rickettsia, which causes a variety of diseases in humans. On the other end of the scale, genetic studies support the endosymbiotic theory according to which mitochondria and related organelles developed from members of this group.[2] Some have also speculated that viruses might have developed from them, or from organisms like them.
The Rickettsiales are among the most mysterious groups of Proteobacteria, owing largely to difficulties in cultivating them. The group includes all obligate endosymbiont bacteria. However, a number of species have been removed, such as Coxiella burnetii, the cause of Q fever.
The Rickettsiales has a sister order the Pelagibacterales, which is composed of free living bacteria with extremely streamlined genomes. Prior to the cultivation of the Pelagibacterales type-species, Pelagibacter ubique, this order was known solely via metagenomic data and was known as SAR11 clade and was a member of the Rickettsiales. The relation between the two order is retained in the subclass, the Rickettsidae, which include the Rickettsiales, the Pelagibacteriales and the extinct protomitochondrion (mitochondria themselves are not bacteria, but organelles).[3]
The Holosporaceae was once a family in the Rickettsiales, but has since been placed in its own order, the Holosporales, which most likely is not as close to the Rickettsiales as initially believed, instead it is a basal order to the sister subclass of the Rickettsidae, the Caulobacteridae.[3]

Phylogeny of Rickettsiales

Magnetococcidae

Magnetococcales

Magnetococcaceae

Magnetococcus marinus

Caulobacteridae

Rhodospirillales, Sphingomonadales, Rhodobacteraceae, Rhizobiales, etc.

Holosporales

Rickettsidae

Pelagibacterales

Pelagibacteraceae

Pelagibacter

subgroups Ib, II, IIIa, IIIb, IV and V

Proto-mitochondria

Anaplasmataceae

Ehrlichia

Anaplasma

Wolbachia

Neorickettsia

분당오피

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